Edward V, King of England - Who actually murdered them?

It is said that when Edward heard how his uncle Richard was crowned in his stead, he sighed, and said, " Ah, would he let me have my life and my freedom he might take the kingdom." After that, in great fear as to what would be their fate, both
the kino; and his brother ;ave themselves over to sorrow and despair. On this particular night they lay sleeping soundly. Tyrrel had with him two men, John Dykewood and Miles Forrest, and these he sent up into the princes' room. In a few minutes they came stealthily down. The deed had been done, and the little princes now slept so soundly that there was no fear of their waking
at the sound of any voice, or further troubling Richard. But Sir James must see for himself, so that he might carry the news to the king. A priest also was summoned, and a place hastily made between the stone staircase, and a hurried funeral took place. Later on the chaplain to Sir John Brackenbury, acting under Richard's orders, had the bodies removed to a more worthy place, but no man knew where he had laid them. It was not for two hundred years that the mystery was cleared up. And then, in the
reign of Charles II., some workmen making alterations found the bodies of two boys. A full inquiry was made, and it was considered certain that these could be none other than the bodies of the little princes, so they were re- moved to Westminster Abbey, and laid at last, where alone was their fitting resting-place, among the kings of England.

Prisoners of the Tower of london Page 81

http://www.searchengine.org.uk/dailyebook/Prisoners%20of%20the%20To...

Wikipedia has this:

Tyrrell was the loyal servant of Richard III who is said to have confessed to the murder of the princes before his execution for treason in 1502. In his history, More said that the princes were smothered to death in their beds by two agents of Tyrrell (Miles Forrest and John Dighton) and were then buried "at the stayre foote, metely depe in the grounde vnder a great heape of stones", but were later disinterred and buried in a secret place. citing this document by Sir Thomas Moore https://scholarsbank.uoregon.edu/xmlui/bitstream/handle/1794/801/ki...

https://en.wikipedia.org/wiki/Princes_in_the_Tower